Error: uniform variables in vertex shader do not fit

On a Nexus 5 device, my game, Ned et les maki, failed with QuadShader related errors:

W/playn (10726): Failed to create QuadShader: java.lang.RuntimeException: Failed to link program: --From Vertex Shader:
W/playn (10726): Error: uniform variables in vertex shader do not fit in 251 vectors.
W/playn (10726): --From Fragment Shader:
W/playn (10726): Error: uniform variables in vertex shader do not fit in 251 vectors.

Full log: http://pastebin.com/JzdC3RzN